About the Authors

Nathan Wiggers

Affiliation Department of Equine Sciences, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Sandra L. P. Nauwelaerts

Affiliation Department of Biology, University of Antwerp, Antwerp, Belgium

Sarah Jane Hobbs

sjhobbs1@uclan.ac.uk

Affiliation Centre for Applied Sport and Exercise Sciences, University of Central Lancashire, Preston, United Kingdom

Sophie Bool

Affiliation Department of Equine Sciences, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Claudia F. Wolschrijn

Affiliation Department of Pathobiology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Willem Back

Affiliations Department of Equine Sciences, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands, Department of Surgery and Anaesthesiology, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Competing Interests

Qualisys AB kindly provided £1000 towards costs and Kistler Instrument Corp loaned equipment FOC. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NW SN SJH SB CW WB. Performed the experiments: NW SN SJH SB CW WB. Analyzed the data: NW SN SJH SB CW WB. Contributed reagents/materials/analysis tools: NW SN SJH SB CW WB. Wrote the paper: NW SN SJH SB CW WB.