About the Authors

Woori Kwak

Contributed equally to this work with: Woori Kwak, Ki-Duk Song

WK and KDS are co-first authors on this work.

Affiliations C & K genomics, Seoul, Republic of Korea, Interdisciplinary Program in Bioinformatics, Seoul National University, Seoul, Republic of Korea

Ki-Duk Song

Contributed equally to this work with: Woori Kwak, Ki-Duk Song

WK and KDS are co-first authors on this work.

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, Republic of Korea

Jae-Don Oh

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, Republic of Korea

Kang-Nyeong Heo

Affiliation Poultry Science Division, National Institute of Animal Science, Cheonan, Republic of Korea

Jun-Heon Lee

Affiliation Department of Animal Science and Biotechnology, Chungnam National University, Daejeon, Republic of Korea

Woon Kyu Lee

Affiliation Inha Research Institute for Medical Sciences, Inha University School of Medicine, Inchon, Republic of Korea

Sook Hee Yoon

Affiliation Department of Agricultural Biotechnology and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, Republic of Korea

Heebal Kim

Affiliation Department of Agricultural Biotechnology and Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, Republic of Korea

Seoae Cho

Affiliation C & K genomics, Seoul, Republic of Korea

Hak-Kyo Lee

breedlee@empas.com

Affiliation Genomic Informatics Center, Hankyong National University, Anseong, Republic of Korea

Competing Interests

Seoae Cho is CEO of C & K genomics, an affiliated company of Seoul National University. Woori Kwak is a current employee of C & K genomics. The remaining authors have no competing interests to report. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KDS HKL. Performed the experiments: JDO. Analyzed the data: WK KNH SHY HK SC. Contributed reagents/materials/analysis tools: JHL WKL. Wrote the paper: WK KDS.