About the Authors

Eriko Kage-Nakadai

Contributed equally to this work with: Eriko Kage-Nakadai, Rieko Imae

Affiliations Department of Physiology, Tokyo Women’s Medical University School of Medicine, Tokyo, Japan, The OCU Advanced Research Institute for Natural Science and Technology, Osaka City University, Osaka, Japan

Rieko Imae

Contributed equally to this work with: Eriko Kage-Nakadai, Rieko Imae

Current address: Graduate School of Pharmaceutical Sciences, The University of Tokyo, Japan

Affiliation Department of Physiology, Tokyo Women’s Medical University School of Medicine, Tokyo, Japan

Yuji Suehiro

Affiliation Department of Physiology, Tokyo Women’s Medical University School of Medicine, Tokyo, Japan

Sawako Yoshina

Affiliation Department of Physiology, Tokyo Women’s Medical University School of Medicine, Tokyo, Japan

Sayaka Hori

Affiliation Department of Physiology, Tokyo Women’s Medical University School of Medicine, Tokyo, Japan

Shohei Mitani

mitani.shohei@twmu.ac.jp

Affiliation Department of Physiology, Tokyo Women’s Medical University School of Medicine, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EKN RI SM. Performed the experiments: EKN RI YS SY SH. Contributed reagents/materials/analysis tools: EKN RI YS SY SH. Contributed to the writing of the manuscript: EKN RI SM.