About the Authors

Yan Liu

Affiliations Department of Microbiology, School of Basic Medical Science, Anhui Medical University, Hefei, Anhui, 230032, China, Department of Pathology, University of Texas Medical Branch, Galveston, Texas, 77555-0609, United States of America

Bin Wu

Affiliation Jiangsu Provincial Center for Diseases Control and Prevention, Nanjing, Jiangsu, China

George Weinstock

Affiliation Department of Genetics, Washington University School of Medicine, St. Louis, MO, 63110, United States of America

David H. Walker

Affiliation Department of Pathology, University of Texas Medical Branch, Galveston, Texas, 77555-0609, United States of America

Xue-jie Yu

xuyu@utmb.edu

Affiliations Department of Pathology, University of Texas Medical Branch, Galveston, Texas, 77555-0609, United States of America, School of Public Health, Shandong University, Jinan, Shandong, 250012, China

Competing Interests

The co-author Xue-jie Yu is a PLOS ONE Editorial Board member, but this does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: XJY GW DHW. Performed the experiments: YL BW. Analyzed the data: XJY YL. Wrote the paper: XJY DHW.