About the Authors

Hussein Khalil

Hussein.khalil@slu.sei

Affiliation Department of Wildlife, Fish, and Environmental Studies, Swedish University of Agricultural Sciences, Umeå, Sweden

Gert Olsson

Affiliation Department of Wildlife, Fish, and Environmental Studies, Swedish University of Agricultural Sciences, Umeå, Sweden

Frauke Ecke

Affiliations Department of Wildlife, Fish, and Environmental Studies, Swedish University of Agricultural Sciences, Umeå, Sweden, Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Sciences, Uppsala, Sweden

Magnus Evander

Affiliation Department of Clinical Microbiology, Division of Virology, Umeå University, Umeå, Sweden

Marika Hjertqvist

Affiliation Swedish Institute for Infectious Disease Control, Stockholm, Sweden

Magnus Magnusson

Affiliation Department of Wildlife, Fish, and Environmental Studies, Swedish University of Agricultural Sciences, Umeå, Sweden

Mikaell Ottosson Löfvenius

Affiliation Department of Forest Ecology and Management, Swedish University of Agricultural Sciences, Umeå, Sweden

Birger Hörnfeldt

Affiliation Department of Wildlife, Fish, and Environmental Studies, Swedish University of Agricultural Sciences, Umeå, Sweden

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BH GO HK MM FE. Performed the experiments: BH. Analyzed the data: HK GO BH. Contributed reagents/materials/analysis tools: BH MH MOL. Wrote the paper: HK GO FE ME MH MM MOL BH.