About the Authors

Nicholas J. Matheson

njm25@cam.ac.uk (NJM); pjl30@cam.ac.uk (PJL)

Affiliation Cambridge Institute for Medical Research, University of Cambridge, Hills Road, Cambridge, United Kingdom

Andrew A. Peden

Affiliation Department of Biomedical Science, University of Sheffield, Western Bank, Sheffield, United Kingdom

Paul J. Lehner

njm25@cam.ac.uk (NJM); pjl30@cam.ac.uk (PJL)

Affiliation Cambridge Institute for Medical Research, University of Cambridge, Hills Road, Cambridge, United Kingdom

Competing Interests

The method described is the subject of UK Patent Application number 1410262.8 on which the authors are named inventors. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: NM AP PL. Performed the experiments: NM. Analyzed the data: NM. Contributed to the writing of the manuscript: NM AP PL.