About the Authors

Li Xu

Affiliation College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China

Zhongyuan Lin

Affiliations College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Jiangsu Key Lab of Marine Biology, Nanjing, China

Qing Tao

Affiliations College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Jiangsu Key Lab of Marine Biology, Nanjing, China

Mingxiang Liang

liangmx@njau.edu.cn

Affiliations College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Jiangsu Key Lab of Marine Biology, Nanjing, China

Gengmao Zhao

Affiliations College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Jiangsu Key Lab of Marine Biology, Nanjing, China

Xiangzhen Yin

Affiliations College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Jiangsu Key Lab of Marine Biology, Nanjing, China

Ruixin Fu

Affiliations College of Resources and Environmental Sciences, Nanjing Agricultural University, Nanjing, China, Jiangsu Key Lab of Marine Biology, Nanjing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ML LX GZ. Performed the experiments: ZL QT LX. Analyzed the data: LX ZL ML. Contributed reagents/materials/analysis tools: RF XY. Contributed to the writing of the manuscript: LX ML ZL.