About the Authors

Yuki Nakazawa

Current address: Center for Neuroscience, University of California Davis, Davis, California, United States of America

Affiliation Department of Biological Sciences, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Tetsuro Ariyoshi

Current address: Department of Neurobiology, Graduate School of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Affiliation Department of Biological Sciences, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Akira Noga

Affiliation Department of Biological Sciences, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Ritsu Kamiya

Affiliations Department of Biological Sciences, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo, Japan, Department of Life Science, Faculty of Science, Gakushuin University, Toshima-ku, Tokyo, Japan

Masafumi Hirono

hirono@bs.s.u-tokyo.ac.jp

Affiliation Department of Biological Sciences, Graduate School of Science, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN RK MH. Performed the experiments: YN TA AN RK MH. Analyzed the data: YN TA AN RK MH. Contributed to the writing of the manuscript: YN RK MH.