About the Authors

Osamu Ichii

ichi-o@vetmed.hokudai.ac.jp

Affiliation Laboratory of Anatomy, Department of Biomedical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Saori Otsuka-Kanazawa

Affiliation Laboratory of Anatomy, Department of Biomedical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Taro Horino

Affiliation Department of Endocrinology, Metabolism and Nephrology, Kochi Medical School, Kochi University, Nankoku, Kochi, Japan

Junpei Kimura

Affiliation Laboratory of Anatomy, Department of Biomedical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Teppei Nakamura

Affiliations Laboratory of Anatomy, Department of Biomedical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan, Section of Biological Safety Research, Chitose Laboratory, Japan Food Research Laboratories, Chitose, Hokkaido, Japan

Manabu Matsumoto

Affiliation Department of Diagnostic Pathology, Kochi Medical School, Kochi University, Nankoku, Kochi, Japan

Makoto Toi

Affiliation Department of Diagnostic Pathology, Kochi Medical School, Kochi University, Nankoku, Kochi, Japan

Yasuhiro Kon

Affiliation Laboratory of Anatomy, Department of Biomedical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

This research was chosen for a TORAY Award (2011) sponsored by Leave a Nest Inc. (Tokyo, Japan). The authors declare that no competing interests exist relating to employment, consultancy, patents, products in development, or marketed products.

Author Contributions

Conceived and designed the experiments: OI SO TH YK. Performed the experiments: OI JK TN. Analyzed the data: OI JK TN. Contributed reagents/materials/analysis tools: TH MM MT. Contributed to the writing of the manuscript: OI SO YK.