About the Authors

Thomas Grönthal

thomas.gronthal@helsinki.fi

Affiliation Central Laboratory, Department of Equine and Small Animal Medicine, University of Helsinki, Helsinki, Finland

Arshnee Moodley

Affiliation Department of Veterinary Disease Biology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Suvi Nykäsenoja

Affiliation Food and Feed Microbiology Research Unit, Finnish Food Safety Authority Evira, Helsinki, Finland

Jouni Junnila

Affiliation 4Pharma Ltd., Espoo, Finland

Luca Guardabassi

Affiliation Department of Veterinary Disease Biology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Katariina Thomson

Affiliation Veterinary Teaching Hospital, University of Helsinki, Helsinki, Finland

Merja Rantala

Affiliation Central Laboratory, Department of Equine and Small Animal Medicine, University of Helsinki, Helsinki, Finland

Competing Interests

JJ is an employee of 4Pharma Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MR TG KT. Performed the experiments: TG MR LG AM SN. Analyzed the data: TG JJ MR. Contributed reagents/materials/analysis tools: AM JJ LG. Wrote the paper: TG AM SN JJ LG KT MR. Molecular analysis: AM SN LG MR TG. Statistical analysis: JJ. Drafted the manuscript: TG.