About the Authors

Kei Higashikawa

Affiliations Graduate School of Medicine, Dentistry, and Pharmaceutical Sciences, Okayama University, Okayama, Japan, Japan Society for the Promotion of Science, Tokyo, Japan

Katsuharu Yagi

Affiliation Graduate School of Medicine, Dentistry, and Pharmaceutical Sciences, Okayama University, Okayama, Japan

Keiko Watanabe

Affiliation Graduate School of Medicine, Dentistry, and Pharmaceutical Sciences, Okayama University, Okayama, Japan

Shinichiro Kamino

Affiliation Next-generation Imaging Team, RIKEN Center for Life Science Technologies, Kobe, Japan

Masashi Ueda

Affiliation Graduate School of Medicine, Dentistry, and Pharmaceutical Sciences, Okayama University, Okayama, Japan

Makoto Hiromura

Affiliation Next-generation Imaging Team, RIKEN Center for Life Science Technologies, Kobe, Japan

Shuichi Enomoto

semo@riken.jp

Affiliations Graduate School of Medicine, Dentistry, and Pharmaceutical Sciences, Okayama University, Okayama, Japan, Next-generation Imaging Team, RIKEN Center for Life Science Technologies, Kobe, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KH KY MH SE. Performed the experiments: KH KY KW. Analyzed the data: KH KY. Contributed reagents/materials/analysis tools: SE. Wrote the paper: KH SK MU MH SE.