About the Authors

Ryan Daly

ryandaly.mail@gmail.com

Affiliations Department of Zoology & Entomology, Rhodes University, Grahamstown, South Africa, Port Elizabeth Museum at Bayworld, Humewood, Port Elizabeth, South Africa

Malcolm J. Smale

Affiliations Port Elizabeth Museum at Bayworld, Humewood, Port Elizabeth, South Africa, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa

Paul D. Cowley

Affiliation South African Institute for Aquatic Biodiversity (SAIAB), Grahamstown, South Africa

Pierre W. Froneman

Affiliation Department of Zoology & Entomology, Rhodes University, Grahamstown, South Africa

Competing Interests

RD and MJS have an affiliation with the institute (Port Elizabeth Museum at Bayworld) that received funding from the National Research Foundation of South Africa for this study. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RD MJS PDC. Performed the experiments: RD. Analyzed the data: RD. Contributed reagents/materials/analysis tools: MJS PDC PWF. Wrote the paper: RD MJS PDC PWF. Obtained permission to conduct the study: RD.