About the Authors

T. E. Anne Cotton

anne.cotton@sheffield.ac.uk

Current address: Department of Animal and Plant Sciences, University of Sheffield, Sheffield, United Kingdom

Affiliations Department of Biology, University of York, York, United Kingdom, School of Biological Sciences, University of Essex, Colchester, United Kingdom

Alex J. Dumbrell

Affiliation School of Biological Sciences, University of Essex, Colchester, United Kingdom

Thorunn Helgason

Affiliation Department of Biology, University of York, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TEAC AJD TH. Performed the experiments: TEAC. Analyzed the data: TEAC AJD TH. Contributed reagents/materials/analysis tools: TEAC AJD TH. Wrote the paper: TEAC AJD TH.