About the Authors

Kenichiro Nishikawa

Affiliation Department of Hematology and Oncology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Naohiro Seo

Affiliation Department of Immuno-Gene Therapy, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Mie Torii

¤ Current address: Kyoto University Center for the Promotion of Interdisciplinary Education and Research, Kyoto, Japan

Affiliation Department of Cellular and Molecular Immunology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Nei Ma

Affiliation Faculty of Health Science, Suzuka University of Medical Science, Suzuka, Mie, Japan

Daisuke Muraoka

Affiliation Department of Immuno-Gene Therapy, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Isao Tawara

Affiliation Department of Hematology and Oncology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Masahiro Masuya

Affiliation Department of Hematology and Oncology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Kyosuke Tanaka

Affiliation Gastroenterology and Hepatology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Yoshiyuki Takei

Affiliation Gastroenterology and Hepatology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Hiroshi Shiku

Affiliation Department of Immuno-Gene Therapy, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Naoyuki Katayama

Contributed equally to this work with: Naoyuki Katayama, Takuma Kato

Affiliation Department of Hematology and Oncology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Takuma Kato

Contributed equally to this work with: Naoyuki Katayama, Takuma Kato

katotaku@doc.medic.mie-u.ac.jp

Affiliation Department of Cellular and Molecular Immunology, Mie University Graduate School of Medicine, Tsu, Mie, Japan

Competing Interests

Co-author Dr. Hiroshi Shiku has served as an editor for this journal and is still currently in that role. The authors confirm that this does not alter their adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: KN TK. Performed the experiments: KN NS TK NM. Analyzed the data: KN TK. Contributed reagents/materials/analysis tools: DM IT MM KT YT MT HS NK. Contributed to the writing of the manuscript: KN TK.