About the Authors

Nicol S. Harper

nicol.harper@eng.ox.ac.uk, nicol.harper@dpag.ox.ac.uk

Current address: Department of Physiology, Anatomy, and Genetics, Sherrington Building, University of Oxford, Oxford, United Kingdom; The Institute of Biomedical Engineering, Department of Engineering Science, Old Road Campus Research Building, University of Oxford, Headington, Oxford, United Kingdom

Affiliations UCL Ear Institute, University College London, London, United Kingdom, Redwood Center for Theoretical Neuroscience, University of California, Berkeley, CA, United States of America

Brian H. Scott

Affiliations Laboratory of Neuropsychology, National Institute of Mental Health, National Institute of Health, Bethesda, MD, United States of America, Center for Neural Science, New York University, New York, NY, United States of America

Malcolm N. Semple

Affiliation Center for Neural Science, New York University, New York, NY, United States of America

David McAlpine

Affiliation UCL Ear Institute, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DM BHS MNS. Performed the experiments: DM BHS. Analyzed the data: NH BHS. Contributed reagents/materials/analysis tools: NH BHS. Wrote the paper: NH.