About the Authors

Chisato Yamamoto

kurousagi0914@gmail.com

Affiliation Graduate School of Fisheries Science and Environmental Studies, Nagasaki University, Nagasaki, Japan

Keisuke Furuta

Affiliation Kobe City Suma Aqualife Park, Kobe, Japan

Michihiro Taki

Affiliation Miyajima Public Aquarium, Hatsukaichi, Hiroshima, Japan

Tadamichi Morisaka

Affiliations Wildlife Research Center of Kyoto University, Sakyo-ku, Kyoto, Japan, Institute of Innovative Science and Technology, Tokai University, Shimizu-ku, Shizuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CY TM. Performed the experiments: CY. Analyzed the data: CY TM. Contributed reagents/materials/analysis tools: MT KF TM. Wrote the paper: CY MT.