About the Authors

William T. Bean

bean@humboldt.edu

Affiliation Humboldt State University, Arcata, California, United States of America

Robert Stafford

Affiliation California Department of Fish and Game, Los Osos, California, United States of America

H. Scott Butterfield

Affiliation The Nature Conservancy, San Francisco, California, United States of America

Justin S. Brashares

Affiliation Department of Environmental Science, Policy and Management, University of California, Berkeley, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WB JB RS HSB. Performed the experiments: WB RS. Analyzed the data: WB. Wrote the paper: WB JB RS HSB.