About the Authors

Ann E. Cooper

Affiliations Department of Small Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America, Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Gavin Herbert Eye Institute, Ophthalmology Research, University of California Irvine, Irvine, California, United States of America

Saija Ahonen

Affiliation Department of Veterinary Biosciences and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland and the Folkhälsan Institute of Genetics, Helsinki, Finland

Jessica S. Rowlan

Affiliations Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Ophthalmology, University of Washington Medical School, Seattle, Washington, United States of America

Alison Duncan

Affiliation Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Eija H. Seppälä

Affiliation Department of Veterinary Biosciences and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland and the Folkhälsan Institute of Genetics, Helsinki, Finland

Päivi Vanhapelto

Affiliation Veterinary Clinic Vetset, Kirkkonummi, Finland

Hannes Lohi

Affiliation Department of Veterinary Biosciences and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland and the Folkhälsan Institute of Genetics, Helsinki, Finland

András M. Komáromy

komaromy@cvm.msu.edu (AMK)

Affiliations Department of Small Animal Clinical Sciences, College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America, Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AEC SA HL AMK. Performed the experiments: AEC SA JSR EHS PV HL AMK. Analyzed the data: AEC SA AD PV HL AMK. Contributed to the writing of the manuscript: AEC SA HL AMK.