About the Authors

Caitlin J. Rinz

Affiliation Department of Genetics and Biochemistry, College of Agriculture, Forestry, and Life Sciences, Clemson University, Clemson, South Carolina, United States of America

Jonathan Levine

Affiliation Department of Small Animal Clinical Sciences, College of Veterinary Medicine and Biomedical Sciences, Texas A&M University, College Station, Texas, United States of America

Katie M. Minor

Affiliation Department of Veterinary and Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Hammon D. Humphries

Affiliation Department of Pathology, School of Medicine, University of California San Diego, La Jolla, California, United States of America

Renee Lara

Affiliation Kingdom Animal Hospital, Bryan, Texas, United States of America

Alison N. Starr-Moss

Affiliation Department of Genetics and Biochemistry, College of Agriculture, Forestry, and Life Sciences, Clemson University, Clemson, South Carolina, United States of America

Ling T. Guo

Affiliation Department of Pathology, School of Medicine, University of California San Diego, La Jolla, California, United States of America

D. Colette Williams

Affiliation R. Prichard Veterinary Medical Teaching Hospital, University of California Davis, Davis, California, United States of America

G. Diane Shelton

gshelton@ucsd.edu (GDS); lclark4@clemson.edu (LAC)

Affiliation Department of Pathology, School of Medicine, University of California San Diego, La Jolla, California, United States of America

Leigh Anne Clark

gshelton@ucsd.edu (GDS); lclark4@clemson.edu (LAC)

Affiliation Department of Genetics and Biochemistry, College of Agriculture, Forestry, and Life Sciences, Clemson University, Clemson, South Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GDS LAC. Performed the experiments: CJR JL HDH LTG DCW. Analyzed the data: CJR JL GDS LAC. Contributed reagents/materials/analysis tools: KMM RL ANSM. Contributed to the writing of the manuscript: CJR ANSM GDS LAC.