About the Authors

Honglong Zhang

Contributed equally to this work with: Honglong Zhang, Zhongjie Li

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Zhongjie Li

Contributed equally to this work with: Honglong Zhang, Zhongjie Li

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Shengjie Lai

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Archie C. A. Clements

Affiliation Research School of Population Health, College of Medicine, Biology and Environment, The Australian National University, Canberra, Australia

Liping Wang

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Wenwu Yin

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Hang Zhou

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Hongjie Yu

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Wenbiao Hu

w.hu@sph.uq.edu.au (WH); yangwz@chinacdc.cn (WY)

Affiliation Infectious Disease Epidemiology Unit, School of Population Health, University of Queensland, Brisbane, Australia

Weizhong Yang

w.hu@sph.uq.edu.au (WH); yangwz@chinacdc.cn (WY)

Affiliation Key Laboratory of Surveillance and Early-warning on infectious Disease, Division of Infectious Diseases, Chinese Center for Disease Control and Prevention, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WZY ZJL. Performed the experiments: HLZ SJL. Analyzed the data: ACAC LPW. Contributed reagents/materials/analysis tools: WWY HZ HJY WBH. Contributed to the writing of the manuscript: ZJL WBH. Designed the steps of analysis: HLZ ZJL.