About the Authors

Catherine E. Thompson

fbsct@leeds.ac.uk

Affiliation University of Leeds, Woodhouse Lane, Leeds, United Kingdom

Jacobus C. Biesmeijer

Affiliation University of Leeds, Woodhouse Lane, Leeds, United Kingdom

Theodore R. Allnutt

Affiliation The Food and Environment Research Agency, Sand Hutton, York, United Kingdom

Stéphane Pietravalle

Affiliation The Food and Environment Research Agency, Sand Hutton, York, United Kingdom

Giles E. Budge

Affiliation The Food and Environment Research Agency, Sand Hutton, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CET GEB TRA. Performed the experiments: CET. Analyzed the data: CET SP. Contributed reagents/materials/analysis tools: CET TRA. Wrote the paper: CET GEB SP JCB.