About the Authors

Keiko Handa

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Kouichi Inukai

kinukai@ks.kyorin-u.ac.jp

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Hirohisa Onuma

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Akihiko Kudo

Affiliation Department of Anatomy, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Fumiyuki Nakagawa

Affiliation Department of Medicine, Shiga University of Medical Science, Otsu, Shiga Japan

Kazue Tsugawa

Affiliation Division of Molecular Biology, Institute for Genome Research, The University of Tokushima, Kuramoto, Tokushima, Japan

Atsuko Kitahara

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Rie Moriya

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Kazuto Takahashi

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Yoshikazu Sumitani

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Toshio Hosaka

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Hayato Kawakami

Affiliation Department of Anatomy, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Seiichi Oyadomari

Affiliation Division of Molecular Biology, Institute for Genome Research, The University of Tokushima, Kuramoto, Tokushima, Japan

Hitoshi Ishida

Affiliation Third Department of Internal Medicine, Division of Diabetes, Endocrinology and Metabolism, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Competing Interests

The authors have no conflicts of interest relevant to this article to declare.

Author Contributions

Conceived and designed the experiments: KI HK SO HI. Performed the experiments: KH AK FN KT HO AK YS. Analyzed the data: RM TH. Contributed to the writing of the manuscript: KI KT.