About the Authors

Zhaoming Qi

Contributed equally to this work with: Zhaoming Qi, Long Huang

Affiliation College of Agriculture, Northeast Agricultural University, Harbin, Heilongjiang, People's Republic of China

Long Huang

Contributed equally to this work with: Zhaoming Qi, Long Huang

Affiliation Biomarker Technologies Corporation, Beijing, China

Rongsheng Zhu

Affiliation College of Agriculture, Northeast Agricultural University, Harbin, Heilongjiang, People's Republic of China

Dawei Xin

Affiliation College of Agriculture, Northeast Agricultural University, Harbin, Heilongjiang, People's Republic of China

Chunyan Liu

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, Heilongjiang, People's Republic of China

Xue Han

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, Heilongjiang, People's Republic of China

Hongwei Jiang

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, Heilongjiang, People's Republic of China

Weiguo Hong

Affiliation Biomarker Technologies Corporation, Beijing, China

Guohua Hu

Affiliation The Crop Research and Breeding Center of Land-Reclamation of Heilongjiang Province, Harbin, Heilongjiang, People's Republic of China

Hongkun Zheng

qshchen@126.com (QC); zhenghk@biomarker.com.cn (HZ)

Affiliation Biomarker Technologies Corporation, Beijing, China

Qingshan Chen

qshchen@126.com (QC); zhenghk@biomarker.com.cn (HZ)

Affiliation College of Agriculture, Northeast Agricultural University, Harbin, Heilongjiang, People's Republic of China

Competing Interests

The authors have read the journal's policy and have the following conflicts: Authors Long Huang, Weiguo Hong and Hongkun Zheng are employed by Biomarker Technologies Corporation, Beijing, China. There are no patents, products in development or marketed products, etc. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ZQ RZ DX QC GH. Performed the experiments: ZQ QC HZ LH. Analyzed the data: LH CL XH HJ. Contributed reagents/materials/analysis tools: ZQ RZ DX CL XH HJ QC GH LH WH HZ. Wrote the paper: ZQ QC WH.