About the Authors

Keisuke Mishiro

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Takahiko Imai

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Sou Sugitani

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Akira Kitashoji

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Yukiya Suzuki

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Toshinori Takagi

Affiliations Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan, Departments of Neurosurgery, Gifu University Graduate School of Medicine, Gifu, Japan

Huayue Chen

Affiliation Department of Anatomy, Gifu University Graduate School of Medicine, Gifu, Japan

Yasunori Oumi

Affiliation Division of Instrument Analysis, Life Science Research Center, Gifu University, Gifu, Japan

Kazuhiro Tsuruma

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Masamitsu Shimazawa

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Hideaki Hara

hidehara@gifu-pu.ac.jp

Affiliation Molecular Pharmacology, Department of Biofunctional Evaluation, Gifu Pharmaceutical University, Gifu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KM KT MS HH TI. Performed the experiments: KM SS AK YS TT HC TI. Analyzed the data: KM SS HC TI. Contributed reagents/materials/analysis tools: TT YO HC. Wrote the paper: KM HC TI.