About the Authors

Tae Heung Kang

kangiron@kku.ac.kr (THK); chung2@jhmi.edu (CFH)

Affiliation Seoul Department of Immunology, College of Medicine, Konkuk University, Chungju, South Korea

Jayne Knoff

Affiliation Department of Pathology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Wei-Hsi Yeh

Affiliation Department of Pathology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Benjamin Yang

Affiliation Department of Pathology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Chenguang Wang

Affiliation Department of Biostatistics and Bioinformatics, Sidney Kimmel Cancer Comprehensive Cancer Center, Johns Hopkins University, Baltimore, Maryland, United States of America

Young Seob Kim

Affiliation Seoul Department of Immunology, College of Medicine, Konkuk University, Chungju, South Korea

Tae Woo Kim

Affiliation Division of Infection and Immunology, Graduate School of Medicine, Korea University, Seoul, South Korea

Tzyy-Choou Wu

Affiliations Department of Pathology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Obstetrics and Gynecology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Molecular Microbiology and Immunology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Oncology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Chien-Fu Hung

kangiron@kku.ac.kr (THK); chung2@jhmi.edu (CFH)

Affiliations Department of Pathology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Oncology, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Competing Interests

TCW is an inventor of intellectual property licensed to Papivax and has ownership interests in Papivax. These interests are being managed by the Johns Hopkins University Conflict of Interest Committee. All other authors declare they have no conflicts of interest. The conflict of interest of author TCW does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: THK TCW CFH. Performed the experiments: THK BY WHY YSK. Analyzed the data: THK TCW CFH TWK CW. Contributed reagents/materials/analysis tools: CFH TCW. Wrote the paper: JK BY WHY TCW CFH.