About the Authors

Ewa Soltysinska

Affiliations The Danish National Research Foundation Centre for Cardiac Arrhythmia, University of Copenhagen, Copenhagen, Denmark, Department of Biomedical Sciences, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Bo Hjorth Bentzen

Affiliation The Danish National Research Foundation Centre for Cardiac Arrhythmia, University of Copenhagen, Copenhagen, Denmark

Maria Barthmes

Affiliations Center for Integrated Protein Science Munich (CIPSM), Ludwig-Maximilians-Universität, Munich, Germany; Department of Pharmacy, Center for Drug Research, Ludwig-Maximilians-Universität, Munich, Germany, Nanion Technologies GmbH, Munich, Germany

Helle Hattel

Affiliation Department of Biomedical Sciences, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

A. Brianne Thrush

Affiliation Department of Biochemistry, Microbiology and Immunology, Faculty of Medicine, University of Ottawa, Ottawa, Canada

Mary-Ellen Harper

Affiliation Department of Biochemistry, Microbiology and Immunology, Faculty of Medicine, University of Ottawa, Ottawa, Canada

Klaus Qvortrup

Affiliation Department of Biomedical Sciences, Core Facility for Integrated Microscopy, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Filip J. Larsen

Affiliation Department of Physiology and Pharmacology, Karolinska Institutet, Stockholm, Sweden

Tomas A. Schiffer

Affiliation Department of Physiology and Pharmacology, Karolinska Institutet, Stockholm, Sweden

Jose Losa-Reyna

Affiliation Department of Physiology and Pharmacology, Karolinska Institutet, Stockholm, Sweden

Julia Straubinger

Affiliation Department of Pharmacology, Toxicology and Clinical Pharmacy, Institute of Pharmacy, University of Tübingen, Tübingen, Germany

Angelina Kniess

Affiliation Department of Pharmacology, Toxicology and Clinical Pharmacy, Institute of Pharmacy, University of Tübingen, Tübingen, Germany

Morten Bækgaard Thomsen

Affiliations The Danish National Research Foundation Centre for Cardiac Arrhythmia, University of Copenhagen, Copenhagen, Denmark, Department of Biomedical Sciences, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Andrea Brüggemann

Affiliation Nanion Technologies GmbH, Munich, Germany

Stefanie Fenske

Affiliation Center for Integrated Protein Science Munich (CIPSM), Ludwig-Maximilians-Universität, Munich, Germany; Department of Pharmacy, Center for Drug Research, Ludwig-Maximilians-Universität, Munich, Germany

Martin Biel

Affiliation Center for Integrated Protein Science Munich (CIPSM), Ludwig-Maximilians-Universität, Munich, Germany; Department of Pharmacy, Center for Drug Research, Ludwig-Maximilians-Universität, Munich, Germany

Peter Ruth

Affiliation Department of Pharmacology, Toxicology and Clinical Pharmacy, Institute of Pharmacy, University of Tübingen, Tübingen, Germany

Christian Wahl-Schott

Affiliation Center for Integrated Protein Science Munich (CIPSM), Ludwig-Maximilians-Universität, Munich, Germany; Department of Pharmacy, Center for Drug Research, Ludwig-Maximilians-Universität, Munich, Germany

Robert Christopher Boushel

Contributed equally to this work with: Robert Christopher Boushel, Søren-Peter Olesen, Robert Lukowski

Affiliation Department of Biomedical Sciences, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Søren-Peter Olesen

Contributed equally to this work with: Robert Christopher Boushel, Søren-Peter Olesen, Robert Lukowski

spfo@sund.ku.dk (SPO); robert.lukowski@uni-tuebingen.de (RL)

Affiliations The Danish National Research Foundation Centre for Cardiac Arrhythmia, University of Copenhagen, Copenhagen, Denmark, Department of Biomedical Sciences, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

Robert Lukowski

Contributed equally to this work with: Robert Christopher Boushel, Søren-Peter Olesen, Robert Lukowski

spfo@sund.ku.dk (SPO); robert.lukowski@uni-tuebingen.de (RL)

Affiliation Department of Pharmacology, Toxicology and Clinical Pharmacy, Institute of Pharmacy, University of Tübingen, Tübingen, Germany

Competing Interests

BHB and SPO have until 2012 worked at and consulted for NeuroSearch that invented NS11021. MB and AB are employed by Nanion Technologies GmbH: This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ES BHB M. Barthmes MBT M. Biel PR CWS RCB SPO RL. Performed the experiments: ES BHB M. Barthmes HH ABT MEH KQ FJL TAS JLR JS AK SF RCB RL. Analyzed the data: ES BHB M. Barthmes MBT M. Biel PR CWS RCB SPO RL. Contributed reagents/materials/analysis tools: AB. Wrote the paper: ES BHB M. Barthmes MBT M. Biel PR CWS RCB RL.