About the Authors

Mieke C. Zwart

Affiliation Newcastle University, School of Biology, Newcastle upon Tyne, United Kingdom

Andrew Baker

Affiliation Baker Consultants Ltd, Matlock, United Kingdom

Philip J. K. McGowan

Contributed equally to this work with: Philip J. K. McGowan, Mark J. Whittingham

Affiliation Newcastle University, School of Biology, Newcastle upon Tyne, United Kingdom

Mark J. Whittingham

Contributed equally to this work with: Philip J. K. McGowan, Mark J. Whittingham

mark.whittingham@newcastle.ac.uk

Affiliation Newcastle University, School of Biology, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist. One of the authors, Andrew Baker, is employed by Baker Consultants Ltd. This does not alter their adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MZ AB PM MW. Performed the experiments: MZ. Analyzed the data: MZ. Contributed reagents/materials/analysis tools: AB. Wrote the paper: MZ AB PM MW.