About the Authors

John M. Marshall


Affiliation Medical Research Council Centre for Outbreak Analysis and Modelling, Department of Infectious Disease Epidemiology, Imperial College London, London, United Kingdom

Bruce A. Hay

Affiliation Division of Biology and Biological Engineering, California Institute of Technology, Pasadena, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JMM BAH. Performed the experiments: JMM. Analyzed the data: JMM. Wrote the paper: JMM BAH.