About the Authors

Nick Wheelhouse

nick.wheelhouse@moredun.ac.uk

Affiliation Moredun Research Institute, Edinburgh, Midlothian, United Kingdom

Christopher Coyle

Affiliation School of Life, Sport and Social Sciences, Edinburgh Napier University, Sighthill Campus, Edinburgh, United Kingdom

Peter G. Barlow

Affiliation School of Life, Sport and Social Sciences, Edinburgh Napier University, Sighthill Campus, Edinburgh, United Kingdom

Stephen Mitchell

Affiliation Institute of Molecular Plant Sciences, University of Edinburgh, Edinburgh, United Kingdom

Gilbert Greub

Affiliation Center for Research on Intracellular Bacteria (CRIB), Institute of Microbiology, University Hospital Center and University of Lausanne, Lausanne, Switzerland

Tim Baszler

Affiliation Washington Animal Disease Diagnostic Laboratory, College of Veterinary Medicine, Washington State University, Pullman, Washington, United States of America

Mick T. Rae

Affiliation School of Life, Sport and Social Sciences, Edinburgh Napier University, Sighthill Campus, Edinburgh, United Kingdom

David Longbottom

Affiliation Moredun Research Institute, Edinburgh, Midlothian, United Kingdom

Competing Interests

Author Gilbert Greub is a PLOS ONE editorial board member. This does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: NW PB GG MR DL. Performed the experiments: NW CC SM. Analyzed the data: NW CC PB MR. Contributed reagents/materials/analysis tools: SM TB GG. Contributed to the writing of the manuscript: NW PB MR GG DL.