About the Authors

Xiaoming Zhang

Contributed equally to this work with: Xiaoming Zhang, Jatupol Kampuansai, Xuebin Qi

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China, Kunming College of Life Science, University of Chinese Academy of Sciences, Beijing, China

Jatupol Kampuansai

Contributed equally to this work with: Xiaoming Zhang, Jatupol Kampuansai, Xuebin Qi

Affiliation Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand

Xuebin Qi

Contributed equally to this work with: Xiaoming Zhang, Jatupol Kampuansai, Xuebin Qi

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China

Shi Yan

Affiliation State Key Laboratory of Genetic Engineering and Ministry of Education, Key Laboratory of Contemporary Anthropology, School of Life Sciences and Institutes of Biomedical Sciences, Fudan University, Shanghai, China

Zhaohui Yang

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China, Kunming College of Life Science, University of Chinese Academy of Sciences, Beijing, China

Bun Serey

Affiliation Department of Geography and Land Management, Royal University of Phnom Penh, Phnom Penh, Cambodia

Tuot Sovannary

Affiliation Department of Geography and Land Management, Royal University of Phnom Penh, Phnom Penh, Cambodia

Long Bunnath

Affiliation Department of Geography and Land Management, Royal University of Phnom Penh, Phnom Penh, Cambodia

Hong Seang Aun

Affiliation Department of Geography and Land Management, Royal University of Phnom Penh, Phnom Penh, Cambodia

Ham Samnom

Affiliation Capacity Development Facilitator for Handicap International Federation and Freelance Research, Battambang, Cambodia

Wibhu Kutanan

Affiliation Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, Thailand

Xin Luo

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China, Kunming College of Life Science, University of Chinese Academy of Sciences, Beijing, China

Shiyu Liao

Affiliations State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China, Kunming College of Life Science, University of Chinese Academy of Sciences, Beijing, China

Daoroong Kangwanpong

Affiliation Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand

Li Jin

Affiliation State Key Laboratory of Genetic Engineering and Ministry of Education, Key Laboratory of Contemporary Anthropology, School of Life Sciences and Institutes of Biomedical Sciences, Fudan University, Shanghai, China

Hong Shi

shihong@mail.kiz.ac.cn (HS); sub@mail.kiz.ac.cn (BS)

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China

Bing Su

shihong@mail.kiz.ac.cn (HS); sub@mail.kiz.ac.cn (BS)

Affiliation State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: B. Su H. Shi. Analyzed the data: XZ JK XQ SY H. Shi B. Su LJ. Wrote the paper: XZ XQ H. Shi B. Su. Collected the samples: XZ JK XQ ZY B. Serey TS LB HSA H. Samnom DK H. Su. Collected the data: XZ JK XQ. Provided technical assistance in the experiments: WK XL SL.