About the Authors

Gregory J. Barord

gjbarord@gmail.com

Affiliations Department of Biology, Graduate Center, City University of New York, New York, New York, United States of America, Department of Biology, Brooklyn College, City University of New York, Brooklyn, New York, United States of America

Frederick Dooley

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Andrew Dunstan

Affiliation School of Biomedical Sciences, University of Queensland, Brisbane St. Lucia, Queensland, Australia

Anthony Ilano

Affiliation Department of Biology, University of San Carlos, Cebu City, Cebu, Philippines

Karen N. Keister

Affiliation Department of Biology, Alaskan Observers Incorporated, Seattle, Washington, United States of America

Heike Neumeister

Affiliation Department of Psychology, Hunter College, City University of New York, New York, New York, United States of America

Thomas Preuss

Affiliation Department of Psychology, Hunter College, City University of New York, New York, New York, United States of America

Shane Schoepfer

Affiliations Department of Biology, University of Washington, Seattle, Washington, United States of America, Department of Earth and Space Sciences, University of Washington, Seattle, Washington, United States of America

Peter D. Ward

Affiliations Department of Biology, University of Washington, Seattle, Washington, United States of America, Department of Earth and Space Sciences, University of Washington, Seattle, Washington, United States of America

Competing Interests

Please note that Karen Keister was employed by Alaskan Observers Incorporated during her work on this project and was entirely unrelated to the research project detailed here. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GJB AD AI PDW. Performed the experiments: GJB AD AI KNK HN TP SS FD PDW. Analyzed the data: GJB KNK FD PDW. Wrote the paper: GJB KNK FD PDW.