About the Authors

Inma Aznar

inma.aznar@ucd.ie

Affiliations Centre for Veterinary Epidemiology and Risk Analysis, School of Veterinary Medicine, University College Dublin, Dublin, Ireland, Quantitative Veterinary Epidemiology group, Wageningen Institute of Animal Sciences, Wageningen University, Wageningen, The Netherlands

Klaas Frankena

Affiliation Quantitative Veterinary Epidemiology group, Wageningen Institute of Animal Sciences, Wageningen University, Wageningen, The Netherlands

Simon J. More

Affiliation Centre for Veterinary Epidemiology and Risk Analysis, School of Veterinary Medicine, University College Dublin, Dublin, Ireland

Clare Whelan

Affiliation Enfer Scientific, Naas, Co. Kildare, Ireland

Wayne Martin

Affiliation Professor Emeritus, University of Guelph, Guelph, Ontario, Canada

Eamonn Gormley

Affiliation School of Veterinary Medicine, University College Dublin, Dublin, Ireland

Leigh A. L. Corner

Affiliation School of Veterinary Medicine, University College Dublin, Dublin, Ireland

Denise Murphy

Affiliation Athlone Regional Veterinary Laboratory, Department of Agriculture, Food and the Marine, Athlone, Ireland

Mart C. M. De Jong

Affiliation Quantitative Veterinary Epidemiology group, Wageningen Institute of Animal Sciences, Wageningen University, Wageningen, The Netherlands

Competing Interests

Clare Whelan is an Enfer employee. Enfer Scientific carried out the assays used in this study. Enfer Group played no role in the study design or the analysis and interpretation of data, nor in the decision to submit the manuscript for publication. Neither Enfer nor its employees played any role that could have inappropriately influenced (biased) this work. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: IA SM KF MDJ WM EG LC DM. Performed the experiments: EG LC DM. Analyzed the data: IA KF MDJ WM. Contributed reagents/materials/analysis tools: IA SM KF MDJ CW WM EG LC DM. Wrote the paper: IA SM KF MDJ CW WM EG LC DM.