About the Authors

Saskia Luttikholt

s.luttikholt@gddiergezondheid.nl

Affiliation Department of Small Ruminant Health, GD Animal Health, Deventer, The Netherlands

Anouk Veldhuis

Affiliation Department of Epidemiology, GD Animal Health, Deventer, The Netherlands

René van den Brom

Affiliation Department of Small Ruminant Health, GD Animal Health, Deventer, The Netherlands

Lammert Moll

Affiliation Department of Small Ruminant Health, GD Animal Health, Deventer, The Netherlands

Karianne Lievaart-Peterson

Affiliation Department of Small Ruminant Health, GD Animal Health, Deventer, The Netherlands

Klaas Peperkamp

Affiliation Department of Pathology, GD Animal Health, Deventer, The Netherlands

Gerdien van Schaik

Affiliation Department of Epidemiology, GD Animal Health, Deventer, The Netherlands

Piet Vellema

Affiliation Department of Small Ruminant Health, GD Animal Health, Deventer, The Netherlands

Competing Interests

All authors are employed by GD Animal Health. The authors affiliation to GD Animal Health does not alter their adherence to PLOS ONE policies on sharing data and materials, within the framework of Dutch privacy legislation.

Author Contributions

Conceived and designed the experiments: SL RV AV GS PV. Performed the experiments: SL RV LM KLP PV. Analyzed the data: LM SL AV. Contributed reagents/materials/analysis tools: LM SL AV. Wrote the paper: SL AV RV LM KLP KP GS PV.