About the Authors

Matthew M. Crane

Contributed equally to this work with: Matthew M. Crane, Ivan B. N. Clark

Affiliation SynthSys - Center for Synthetic and Systems Biology and School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Ivan B. N. Clark

Contributed equally to this work with: Matthew M. Crane, Ivan B. N. Clark

Affiliation SynthSys - Center for Synthetic and Systems Biology and School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Elco Bakker

Affiliation SynthSys - Center for Synthetic and Systems Biology and School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Stewart Smith

Affiliation Scottish Microelectronics Center, University of Edinburgh, Edinburgh, United Kingdom

Peter S. Swain

peter.swain@ed.ac.uk

Affiliation SynthSys - Center for Synthetic and Systems Biology and School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MMC IBNC EB SS PSS. Performed the experiments: MMC IBNC EB. Analyzed the data: MMC IBNC EB SS PSS. Wrote the paper: MMC IBNC EB PSS.