About the Authors

Ruohan Wang

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory for Genetics and Breeding of Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University,Beijing, China

Sai Xu

Affiliations National Engineering Laboratory for Tree Breeding, Key Laboratory for Genetics and Breeding of Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University,Beijing, China, School of Environment, Tsinghua University, Beijing, China

Xiangyu Liu

Affiliation Lab of Systematic Evolution and Biogeography of Woody Plants, College of Nature Conservation, Beijing Forestry University,Beijing, China

Yiyuan Zhang

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory for Genetics and Breeding of Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University,Beijing, China

Jianzhong Wang

Affiliation National Engineering Laboratory for Tree Breeding, Key Laboratory for Genetics and Breeding of Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Biotechnology, Beijing Forestry University,Beijing, China

Zhixiang Zhang

zxzhang@bjfu.edu.cn

Affiliation Lab of Systematic Evolution and Biogeography of Woody Plants, College of Nature Conservation, Beijing Forestry University,Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RW ZZ JW. Performed the experiments: RW XL SX YZ. Analyzed the data: RW SX ZZ. Contributed reagents/materials/analysis tools: SX YZ. Wrote the paper: RW SX XL ZZ. Provided crucial suggestions on the experiment: ZZ JW.