About the Authors

Lucas D. Tilley

Affiliation Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America

Shruthi Krishnamurthy

Affiliation Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America

Nicholas J. Westwood

Affiliation School of Chemistry and Biomedical Sciences Research Complex, University of St Andrews and EaStCHEM, North Haugh, St Andrews, Fife, Scotland, United Kingdom

Gary E. Ward

Gary.Ward@uvm.edu

Affiliation Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America

Competing Interests

Gary E. Ward is a member of the PLOS Board of Directors. This does not alter the authors' adherence to PLOS ONE's policies on sharing data and materials. The authors have declared that no other competing interests exist.

Author Contributions

Conceived and designed the experiments: LDT GEW. Performed the experiments: LDT SK. Analyzed the data: LDT GEW SK. Contributed reagents/materials/analysis tools: LDT NJW GEW. Wrote the paper: LDT GEW.