About the Authors

Kimberly B. Zumbrennen-Bullough

Affiliation Program in Anemia Signaling Research, Division of Nephrology, Program in Membrane Biology, Center for Systems Biology, Massachusetts General Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

Lore Becker

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Experimental Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Lillian Garrett

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Development Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Sabine M. Hölter

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Development Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Julia Calzada-Wack

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Pathology, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Ilona Mossbrugger

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Pathology, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Leticia Quintanilla-Fend

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Pathology, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Ildiko Racz

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Molecular Psychiatry, Life & Brain Center, University of Bonn, Bonn, Germany

Birgit Rathkolb

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Molecular Animal Breeding and Biotechnology, Gene Center, Ludwig-Maximilians-Universitat, Munich, Germany

Thomas Klopstock

Affiliations Department of Neurology, Friedrich-Baur-Institute, Klinikum der Ludwig-Maximilians-Universitat, Munich, Germany, Deutsches Zentrum für Neurodegenerative Erkrankungen, Munich, Germany, Munich Cluster for Systems Neurology, Munich, Germany

Wolfgang Wurst

Affiliations Institute of Development Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Chair of Developmental Genetics, Technische Universitat München, Freising-Weihenstephan, Germany, Max Planck Institute of Psychiatry, Munich, Germany, Deutsches Zentrum für Neurodegenerative Erkrankungen, Munich, Germany, Munich Cluster for Systems Neurology, Munich, Germany

Andreas Zimmer

Affiliation Institute of Molecular Psychiatry, Life & Brain Center, University of Bonn, Bonn, Germany

Eckhard Wolf

Affiliation Institute of Molecular Animal Breeding and Biotechnology, Gene Center, Ludwig-Maximilians-Universitat, Munich, Germany

Helmut Fuchs

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Experimental Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Valerie Gailus-Durner

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Experimental Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany

Martin Hrabě de Angelis

Affiliations German Mouse Clinic, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Institute of Experimental Genetics, Helmholtz-Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany, Chair of Experimental Genetics, Center of Life and Food Sciences Weihenstephan, Technische Universitat München, Freising, Germany, German Center for Diabetes Research, Neuherberg, Germany

Steven J. Romney

Affiliation University of Utah, Department of Medicine, Division of Hematology and Hematological Malignancies, Salt Lake City, Utah, United States of America

Elizabeth A. Leibold

betty.leibold@genetics.utah.edu

Affiliation University of Utah, Department of Medicine, Division of Hematology and Hematological Malignancies, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EAL KZB MHA VGD HF SMH WW. Performed the experiments: KZB SJR BR LB. Analyzed the data: EAL KZB SJR BR EW LG SMH LB IM JCW LQF IR TK. Wrote the paper: EAL KZB. Reviewed the manuscript: TK EW AZ.