About the Authors

Anke Bill

Contributed equally to this work with: Anke Bill, Elizabeth M. Rosethorne

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Elizabeth M. Rosethorne

Contributed equally to this work with: Anke Bill, Elizabeth M. Rosethorne

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

Toby C. Kent

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

Lindsay Fawcett

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

Lynn Burchell

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

Michiel T. van Diepen

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

Anthony Marelli

Affiliation Infectious Diseases, Genomics Institute of the Novartis Research Foundation, San Diego, California, United States of America

Sergey Batalov

Affiliation Infectious Diseases, Genomics Institute of the Novartis Research Foundation, San Diego, California, United States of America

Loren Miraglia

Affiliation Infectious Diseases, Genomics Institute of the Novartis Research Foundation, San Diego, California, United States of America

Anthony P. Orth

Affiliation Infectious Diseases, Genomics Institute of the Novartis Research Foundation, San Diego, California, United States of America

Nicole A. Renaud

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Steven J. Charlton

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

Martin Gosling

Affiliation Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom

L. Alex Gaither

Affiliation Developmental and Molecular Pathways, Novartis Institutes for Biomedical Research, Cambridge, Massachusetts, United States of America

Paul J. Groot-Kormelink

paul.groot-kormelink@novartis.com

Affiliations Respiratory Disease Area, Novartis Institutes for Biomedical Research, Horsham, United Kingdom, Musculoskeletal Disease Area, Novartis Institutes for Biomedical Research, Basel, Switzerland

Competing Interests

All authors are employees of Novartis Institutes for Biomedical Research. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AB ER TCK APO LM SJC MG LAG PJGK. Performed the experiments: AB ER TCK LF LB MTvD AM SB NAR. Analyzed the data: AB ER TCK LF LB MTvD AM SB NAR. Wrote the paper: AB ER TCK APO LM SJC MG LAG PJGK.