About the Authors

Sean C. P. Coogan

scoogan@ualberta.ca

Affiliation Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada

David Raubenheimer

Affiliation The Charles Perkins Centre, Faculty of Veterinary Science, and School of Biological Science, University of Sydney, Sydney, Australia

Gordon B. Stenhouse

Affiliation Grizzly Bear Program, Foothills Research Institute, Hinton, AB, Canada

Scott E. Nielsen

Affiliation Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SCPC DR SEN. Performed the experiments: SCPC DR. Analyzed the data: SCPC DR. Contributed reagents/materials/analysis tools: SEN GBS. Wrote the paper: SCPC. Critically reviewed and revised the manuscript: DR GBS SEN.