About the Authors

Katie Nightingale

Affiliation The Roslin Institute & R(D)SVS, University of Edinburgh, Edinburgh, Midlothian, United Kingdom

Claire S. Levy

Current address: The Scripps Research Institute, La Jolla, California, United States of America

Affiliation The Roslin Institute & R(D)SVS, University of Edinburgh, Edinburgh, Midlothian, United Kingdom

John Hopkins

Affiliation The Roslin Institute & R(D)SVS, University of Edinburgh, Edinburgh, Midlothian, United Kingdom

Finn Grey

Affiliation The Roslin Institute & R(D)SVS, University of Edinburgh, Edinburgh, Midlothian, United Kingdom

Suzanne Esper

Affiliation The Roslin Institute & R(D)SVS, University of Edinburgh, Edinburgh, Midlothian, United Kingdom

Robert G. Dalziel

bob.dalziel@roslin.ed.ac.uk

Affiliation The Roslin Institute & R(D)SVS, University of Edinburgh, Edinburgh, Midlothian, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KN RGD JH FG. Performed the experiments: KN CSL SE. Analyzed the data: KN JH FG RGD. Wrote the paper: RGD.