About the Authors

Bradley E. Carlson

bec169@psu.edu

Affiliations Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Intercollege Graduate Degree Program in Ecology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Shannen McGinley

Affiliation Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Matthew P. Rowe

Current address: Department of Zoology, Michigan State University, East Lansing, Michigan, United States of America

Affiliation Department of Biological Sciences, Sam Houston State University, Huntsville, Texas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BEC SM MPR. Performed the experiments: BEC SM MPR. Analyzed the data: BEC. Contributed reagents/materials/analysis tools: MPR. Wrote the paper: BEC SM MPR.