About the Authors

Earl E. Werner

eewerner@umich.edu

Affiliation Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, United States of America

Christopher J. Davis

Affiliation Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, Michigan, United States of America

David K. Skelly

Affiliation School of Forestry & Environmental Studies, Yale University, New Haven, Connecticut, United States of America

Rick A. Relyea

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Michael F. Benard

Affiliation Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America

Shannon J. McCauley

Affiliation Department of Biology, University of Toronto Mississauga, Mississauga, Ontario, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EW CD DS RR MB SM. Performed the experiments: EW CD DS RR MB SM. Analyzed the data: EW CD DS RR MB SM. Wrote the paper: EW CD DS RR MB SM. Processed samples: CD.