About the Authors

Richard G. Smith

richard.smith@unh.edu

Affiliation Department of Natural Resources and the Environment, University of New Hampshire, Durham, New Hampshire, United States of America

Lesley W. Atwood

Affiliation Department of Natural Resources and the Environment, University of New Hampshire, Durham, New Hampshire, United States of America

Nicholas D. Warren

Affiliation Department of Natural Resources and the Environment, University of New Hampshire, Durham, New Hampshire, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RGS. Performed the experiments: RGS LWA NDW. Analyzed the data: RGS LWA NDW. Contributed reagents/materials/analysis tools: RGS. Wrote the paper: RGS LWA NDW.