About the Authors

Bin Zheng

Affiliation Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands

Jeroen van Bergenhenegouwen

Affiliations Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands, Nutricia Research, Utrecht, The Netherlands

Saskia Overbeek

Affiliation Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands

Hendrik J. G. van de Kant

Affiliation Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands

Johan Garssen

Affiliations Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands, Nutricia Research, Utrecht, The Netherlands

Gert Folkerts

Affiliation Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands

Paul Vos

Affiliation Nutricia Research, Utrecht, The Netherlands

Mary E. Morgan

Affiliation Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands

Aletta D. Kraneveld

A.D.Kraneveld@uu.nl

Affiliation Division of Pharmacology, Utrecht Institute for Pharmaceutical Science, Faculty of Science, Utrecht University, Utrecht, The Netherlands

Competing Interests

Prof. Johan Garssen, besides being employed as a professor at Utrecht University, is also employed by Danone Research B.V. as a research manager. The research presented in this manuscript, however, was performed independently of his Danone Research B.V. duties. Paul Vos and Jeroen van Bergenhenegouwen are also employed by Danone Research, however they were not directly involved in the analysis of the data. The authors declare that this does not alter their adherence to all the PLOS ONE policies on sharing data and materials. The remaining authors have no competing interests.

Author Contributions

Conceived and designed the experiments: BZ MEM ADK. Performed the experiments: BZ MEM JvB HJGvdK SO. Analyzed the data: BZ MEM ADK GF JG. Contributed reagents/materials/analysis tools: HJGvdK SO JvB PV. Wrote the paper: BZ MEM ADK GF JG.