About the Authors

Junjie Luo

Contributed equally to this work with: Junjie Luo, Zhiyuan Song

Affiliation State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China

Zhiyuan Song

Contributed equally to this work with: Junjie Luo, Zhiyuan Song

Affiliation State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China

Shengli Yu

Affiliation State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China

Dan Cui

Affiliation State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China

Benli Wang

Affiliation State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China

Fangrong Ding

Affiliation Beijing GeneProtein Biotechnology Co., Ltd., Beijing, People’s Republic of China

Song Li

Affiliation Beijing GeneProtein Biotechnology Co., Ltd., Beijing, People’s Republic of China

Yunping Dai

daiyunping@sina.com (YPD); ninglcau@cau.edu.cn (NL)

Affiliations State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China, Beijing GeneProtein Biotechnology Co., Ltd., Beijing, People’s Republic of China

Ning Li

daiyunping@sina.com (YPD); ninglcau@cau.edu.cn (NL)

Affiliations State Key Laboratory for Agrobiotechnology, College of Biological Science, China Agricultural University, Beijing, People’s Republic of China, College of Animal Science, Yunnan Agricultural University, Kunming, People’s Republic of China

Competing Interests

YPD, FRD and SL are employees of Beijing Gene Protein Biotechnology Co., Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JJL ZYS SLY YPD NL. Performed the experiments: JJL ZYS SLY BLW DC FRD SL. Analyzed the data: JJL ZYS. Contributed reagents/materials/analysis tools: JJL ZYS SLY YPD NL. Wrote the paper: JJL ZYS NL.