About the Authors

Xiaoting Zheng

Affiliations Institute of Geology and Paleontology, Linyi University, Linyi, Shandong, China, Tianyu Natural History Museum of Shandong, Pingyi, Shandong, China

Jingmai K. O'Connor

jingmai.oconnor@gmail.com

Affiliation Key Laboratory of Vertebrate Evolution and Human Origin of the Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Beijing, China

Fritz Huchzermeyer

Affiliation Department of Paraclinical Sciences, Faculty of Veterinary Science, University of Pretoria, Onderstepoort, South Africa

Xiaoli Wang

Affiliation Institute of Geology and Paleontology, Linyi University, Linyi, Shandong, China

Yan Wang

Affiliation Institute of Geology and Paleontology, Linyi University, Linyi, Shandong, China

Xiaomei Zhang

Affiliation Tianyu Natural History Museum of Shandong, Pingyi, Shandong, China

Zhonghe Zhou

Affiliation Key Laboratory of Vertebrate Evolution and Human Origin of the Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

† Deceased.

Conceived and designed the experiments: X. Zheng ZZ JO. Performed the experiments: JO ZZ XW X. Zhang. Analyzed the data: JO ZZ FH XW YW. Contributed reagents/materials/analysis tools: X. Zheng. Wrote the paper: JO FH ZZ.