About the Authors

Kazuki Morioka

Affiliation Exotic Disease Research Station, National Institute of Animal Health, National Agriculture and Food Research Organization, Josuihoncho Kodaira, Tokyo, Japan

Katsuhiko Fukai

Affiliation Exotic Disease Research Station, National Institute of Animal Health, National Agriculture and Food Research Organization, Josuihoncho Kodaira, Tokyo, Japan

Kenichi Sakamoto

Affiliation Exotic Disease Research Station, National Institute of Animal Health, National Agriculture and Food Research Organization, Josuihoncho Kodaira, Tokyo, Japan

Kazuo Yoshida

Affiliation Exotic Disease Research Station, National Institute of Animal Health, National Agriculture and Food Research Organization, Josuihoncho Kodaira, Tokyo, Japan

Toru Kanno

kannot@affrc.go.jp

Affiliation Exotic Disease Research Station, National Institute of Animal Health, National Agriculture and Food Research Organization, Josuihoncho Kodaira, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KM KY. Performed the experiments: KM KF KY. Analyzed the data: KM. Contributed reagents/materials/analysis tools: KM KF KY TK. Wrote the paper: KM KF KS KY TK.