About the Authors

Erwin Reiling

Affiliations National Institute for Public Health and the Environment, Bilthoven, The Netherlands, Department of Cell Biology and Genetics, Center for Biomedical Genetics, Erasmus MC, Rotterdam, The Netherlands

Martijn E. T. Dollé

Affiliation National Institute for Public Health and the Environment, Bilthoven, The Netherlands

Sameh A. Youssef

Affiliation Faculty of Veterinary Medicine, Department of Pathobiology, Utrecht University, Utrecht, The Netherlands

Moonsook Lee

Affiliation Department of Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Bhawani Nagarajah

Affiliation National Institute for Public Health and the Environment, Bilthoven, The Netherlands

Marianne Roodbergen

Affiliation National Institute for Public Health and the Environment, Bilthoven, The Netherlands

Piet de With

Affiliation National Institute for Public Health and the Environment, Bilthoven, The Netherlands

Alain de Bruin

Affiliation Faculty of Veterinary Medicine, Department of Pathobiology, Utrecht University, Utrecht, The Netherlands

Jan H. Hoeijmakers

Affiliation Department of Cell Biology and Genetics, Center for Biomedical Genetics, Erasmus MC, Rotterdam, The Netherlands

Jan Vijg

Affiliation Department of Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Harry van Steeg

Affiliations National Institute for Public Health and the Environment, Bilthoven, The Netherlands, Department of Toxicogenetics, Leiden University Medical Center, Leiden, The Netherlands

Paul Hasty

hastye@uthscsa.edu

Affiliation Department of Molecular Medicine and Institute of Biotechnology, Barshop Institute for Longevity and Aging Studies, Cancer Therapy and Research Center, University of Texas Health Science Center at San Antonio, San Antonio, Texas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ER METD JHH JV HVS PH. Performed the experiments: ER SAY ML BN MR PDW ADB. Analyzed the data: ER METD JHH JV HVS PH. Wrote the paper: ER PH.