About the Authors

Patrick I. Chiyo

Affiliation Department of Biological Sciences, University of Notre Dame, Notre Dame, Indiana, United States of America

Laura E. Grieneisen

Affiliation Department of Biological Sciences, University of Notre Dame, Notre Dame, Indiana, United States of America

George Wittemyer

Affiliations Department of Fish, Wildlife and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America, Save the Elephants, Nairobi, Kenya

Cynthia J. Moss

Affiliation Amboseli Trust for Elephants, Langata, Nairobi, Kenya

Phyllis C. Lee

Affiliations Amboseli Trust for Elephants, Langata, Nairobi, Kenya, Behaviour and Evolution Research Group, Department of Psychology, University of Stirling, Scotland, United Kingdom

Iain Douglas-Hamilton

Affiliation Save the Elephants, Nairobi, Kenya

Elizabeth A. Archie

earchie@nd.edu

Affiliation Department of Biological Sciences, University of Notre Dame, Notre Dame, Indiana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PIC GW EAA. Performed the experiments: PIC EAA GW CJM PCL ID. Analyzed the data: PIC LG. Contributed reagents/materials/analysis tools: GW CJM PCL ID. Wrote the paper: PIC EAA.