About the Authors

Robert J. Dusek

rdusek@usgs.gov

Affiliation National Wildlife Health Center, United States Geological Survey, Madison, Wisconsin, United States of America

Gunnar T. Hallgrimsson

Current address: Institute of Life and Environmental Sciences, University of Iceland, Reykjavik, Iceland

Affiliation Southwest Iceland Nature Research Institute, Sandgerði, Iceland

Hon S. Ip

Affiliation National Wildlife Health Center, United States Geological Survey, Madison, Wisconsin, United States of America

Jón E. Jónsson

Affiliation Snæfellsnes Research Centre, University of Iceland, Stykkishólmur, Iceland

Srinand Sreevatsan

Affiliation Veterinary and Biomedical Sciences Department, College of Veterinary Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America

Sean W. Nashold

Affiliation National Wildlife Health Center, United States Geological Survey, Madison, Wisconsin, United States of America

Joshua L. TeSlaa

Affiliation National Wildlife Health Center, United States Geological Survey, Madison, Wisconsin, United States of America

Shinichiro Enomoto

Current address: Department of Biology, University of Utah, Salt Lake City, Utah, United States of America

Affiliation Veterinary and Biomedical Sciences Department, College of Veterinary Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America

Rebecca A. Halpin

Affiliation J. Craig Venter Institute, Rockville, Maryland, United States of America

Xudong Lin

Affiliation J. Craig Venter Institute, Rockville, Maryland, United States of America

Nadia Fedorova

Affiliation J. Craig Venter Institute, Rockville, Maryland, United States of America

Timothy B. Stockwell

Affiliation J. Craig Venter Institute, Rockville, Maryland, United States of America

Vivien G. Dugan

Current address: Division of Microbiology and Infectious Diseases, National Institutes of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Affiliation J. Craig Venter Institute, Rockville, Maryland, United States of America

David E. Wentworth

Affiliation J. Craig Venter Institute, Rockville, Maryland, United States of America

Jeffrey S. Hall

Affiliation National Wildlife Health Center, United States Geological Survey, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJD JSH GTH. Performed the experiments: JSH DEW VGD SS RJD JEJ GTH SWN JLT RAH XL TBS NF SE. Analyzed the data: RJD HSI. Wrote the paper: RJD JSH HSI.